Brachinus

Brachinus
	Brachinus crepitans
	Nomenklatur
	Brachinus; Weber, 1801
	Klassifikasjon
Rike	Dyr
Rekke	Leddyr
Klasse	Insekter
Orden	Biller
Familie	Løpebiller
Underfamilie	Bombardérbiller
	Økologi
Antall arter:	ca. 320, ca. 40 i Europa
Habitat:	på land
Utbredelse:	den palearktiske sone, Det nære østen og Nord-Afrika, ikke i Norge
	Inndelt i
	se artsliste;

Brachinus er en slekt av billene i delgruppen (underfamilien) bombardérbiller (Brachininae) i familiegruppen løpebiller. Ingen arter er registret fra Norge.

Utseende

Små til middelsstore (gjerne 5-10 millimeter), spinkle løpebiller, dekkvingene forholdsvis brede. På farge er dekkvingene ofest mørke, ofte med metallglans, hode, pronotum og bein ofte rødlige. Hodet er omtrent femkantet, fasettøynene forholdsvis små, antennene lange og tynne. Pronotum er omtrent så bredt som hodet, bredere foran enn bak, med innsvungne kanter. Dekkvingene er noe flate, oftest med fine punktstriper.

Levevis

Artene lever på varme, tørre, solåpne steder. De voksne billene er rovdyr og kan noen ganger finnes tallrikt under steiner. Larvene er ektoparasitter, i alle fall noen av artene angriper pupper av andre biller. Billene i denne slekten forsvarer seg ved å sprute kokvarme kjemikalier mot angripere, dette kan de gjøre med stor presisjon.

Utbredelse

Slekten har en holarktisk utbredelse, inkludert det meste av Europa, men er ikke funnet i Norge. Det er tallrike arter i Middelhavsområdet.

Systematisk inndeling

Treliste

Ordenen biller, Coleoptera Linnaeus, 1758
- Underordenen Adephaga Schellenberg, 1806
  - Overfamilien løpebiller og vannkalver, Caraboidea Latreille, 1802
    - Familien løpebiller, Carabidae Latreille, 1802
      - Underfamilien bombardérbiller, Brachininae Bonelli, 1810
        Slekten Brachinus Weber, 1801 – tallrike arter i det sørlige Europa

Liste over artene

Brachinus aabaaba Erwin, 1970
Brachinus abbreviatus (Castelnau, 1835)
Brachinus abyssinicus Chaudoir, 1876
Brachinus adustipennis Erwin, 1969
Brachinus aeger Chaudoir, 1876
Brachinus aeneicostis Bates, 1883
Brachinus afghanus Jedlicka, 1967
Brachinus agraphus (Alluaud, 1899)
Brachinus albarracinus Wagner, 1926
Brachinus alexandri F. Battoni, 1984
Brachinus alexiguus Erwin, 1970
Brachinus algoensis Peringuey, 1896
Brachinus alternans Dejean, 1825
Brachinus americanus (Leconte, 1844)
Brachinus andalusiacus Rambur, 1837
Brachinus andreaei Basilewsky, 1964
Brachinus angustatus Dejean, 1831
Brachinus ankarensis Jedlicka, 1962
Brachinus annulicornis Chaudoir, 1842
Brachinus apicalis Erichson, 1843
Brachinus arboreus Chevrolat, 1834
Brachinus aristokrates Liebke, 1934
Brachinus armiger Dejean, 1831
Brachinus atramentarius Mannerheim, 1837
Brachinus atripennis Ballion, 1871
Brachinus azureipennis Chaudoir, 1876
Brachinus baeticus Rambur, 1837
Brachinus bagdatensis Pic, 1902
Brachinus barbarus Lucas, 1846
Brachinus bayardi Dejean, 1831
Brachinus bellicosus L. Dufour, 1820
Brachinus berytensis Reiche & Saulcy, 1855
Brachinus bigutticeps Chaudoir, 1876
Brachinus bilineatus Castelnau, 1835
Brachinus bipustulatus Quensel, 1806
Brachinus bodemeyeri Apfelbeck, 1904
Brachinus brevicollis Motschulsky, 1844
Brachinus brittoni Ali, 1967
Brachinus bruchi Liebke, 1939
Brachinus brunneus Castelnau, 1834
Brachinus brunniventris Motschulsky, 1864
Brachinus caffer Boheman, 1848
Brachinus canaliculatus Fairmaire, 1897
Brachinus capnicus Erwin, 1970
Brachinus catalonicus Jeanne, 1988
Brachinus chalchihuitlicue Erwin, 1970
Brachinus charis Andrewes, 1923
Brachinus chaudoirianus Jakobson, 1908
Brachinus chinensis Chaudoir, 1850
Brachinus chirriador Erwin, 1970
Brachinus chuji Jedlicka, 1956
Brachinus cibolensis Erwin, 1970
Brachinus ciliatus Liebke, 1933
Brachinus cinctellus Chaudoir, 1876
Brachinus cinctipennis Chevrolat, 1835
Brachinus circumtinctus Bates, 1892
Brachinus collarti Basilewsky, 1948
Brachinus conformis Dejean, 1831
Brachinus congicus (Basilewsky, 1958)
Brachinus connectoides (Jeannel, 1949)
Brachinus connectus Dejean, 1831
Brachinus consanguineus Chaudoir, 1876
Brachinus constrictus Reitter, 1919
Brachinus cordicollis Dejean, 1826
Brachinus costatulus Quensel, 1806
Brachinus costiger Chaudoir, 1876
Brachinus costipennis Motschulsky, 1859
Brachinus costulipennis Liebke, 1928
Brachinus crepitans Linne, 1758 – kjent fra Sverige, Finland og Danmark
Brachinus cruciatus Quensel, 1806
Brachinus cyanipennis Say, 1823
Brachinus cyanochroaticus Erwin, 1969
Brachinus daliensis Kirschenhofer, 2011
Brachinus dawnaensis Kirschenhofer, 2003
Brachinus demoulini Basilewsky, 1962
Brachinus dieganus (Jeannel, 1949)
Brachinus diffusus Chaudoir, 1876
Brachinus dilapsus Antoine, 1941
Brachinus dilatatus Klug, 1832
Brachinus dilatipennis Reitter, 1919
Brachinus dilottii Maran, 1933
Brachinus dissimilis (Basilewsky, 1943)
Brachinus distans Lorenz, 1998
Brachinus dorsalis Dejean, 1831
Brachinus dryas Andrewes, 1936
Brachinus dryas Andrewes, 1936
Brachinus efflans Dejean, 1830
Brachinus ejaculans Fischer von Waldheim, 1828
Brachinus elegans Chaudoir, 1842
Brachinus elegantulus Erichson, 1842
Brachinus elongatulus Chaudoir, 1876
Brachinus enonensis Peringuey, 1899
Brachinus eucosmus Andrewes, 1937
Brachinus evanescens Bates, 1892
Brachinus exhalans P. Rossi, 1792
Brachinus explodens Duftschmid, 1812
Brachinus explosus Erwin, 1970
Brachinus exquisitus Bates, 1892
Brachinus fageli Basilewsky, 1962
Brachinus fasciatocollis (Fairmaire, 1901)
Brachinus favicollis Erwin, 1965
Brachinus femoralis Gistel, 1857
Brachinus flavicapillus Bates, 1892
Brachinus flaviventris Chaudoir, 1876
Brachinus flavus Jedlicka, 1955
Brachinus foochowi Kirschenhofer, 1986
Brachinus formosanus Jedlicka, 1939
Brachinus forticostis Liebke, 1934
Brachinus frontalis Chaudoir In Oberthur, 1883
Brachinus fulminatus Erwin, 1969
Brachinus fulvipennis Chaudoir, 1876
Brachinus fumans (Fabricius, 1781)
Brachinus fumator Gistel, 1857
Brachinus fuscicornis Dejean, 1826
Brachinus fuscipennis Dejean, 1825
Brachinus galactoderus Erwin, 1970
Brachinus gebhardis Erwin, 1965
Brachinus genicularis Mannerheim, 1837
Brachinus geniculatus Dejean, 1831
Brachinus gentilis Erichson, 1843
Brachinus ghindanus Liebke, 1934
Brachinus grandis Brulle, 1838
Brachinus hamatus Fischer von Waldheim, 1828
Brachinus hazardi Andrewes, 1930
Brachinus hexagrammus Chaudoir, 1876
Brachinus hirsutus Bates, 1884
Brachinus hoffmanni Liebke, 1927
Brachinus humeralis Ahrens, 1812
Brachinus hylaenus Reichardt, 1967
Brachinus ichabodopsis Erwin, 1970
Brachinus illotus Chaudoir, 1876
Brachinus immaculicornis Dejean, 1826
Brachinus immarginatus Brulle, 1838
Brachinus imperialensis Erwin, 1965
Brachinus imporcitis Erwin, 1970
Brachinus incomptus Bates, 1873
Brachinus inconditus Peringuey, 1896
Brachinus inops Andrewes, 1932
Brachinus intactus Bates, 1892
Brachinus intermedius Brulle, 1838
Brachinus irakus Liebke, 1933
Brachinus italicus Dejean, 1831
Brachinus janthinipennis (Dejean, 1831)
Brachinus javalinopsis Erwin, 1970
Brachinus jucundus Dejean, 1831
Brachinus kalalovae Roubal, 1932
Brachinus kansanus Leconte, 1862
Brachinus kavanaughi Erwin, 1969
Brachinus knirschi Jedlicka, 1931
Brachinus kollari Liebke, 1934
Brachinus krynickii Jedlick
Brachinus kryzhanovskii Belousov & Kabak, 1992
Brachinus laetus Dejean, 1831
Brachinus laevicostis Liebke, 1934
Brachinus langenhani Liebke, 1934
Brachinus lapidarius Basilewsky, 1962
Brachinus lateralis Dejean, 1831
Brachinus latipennis Peyerimhoff, 1907
Brachinus latus Lorenz, 1998
Brachinus lavaudeni (Jeannel, 1949)
Brachinus leprieuri Gory, 1833
Brachinus lesnei Andrewes, 1923
Brachinus lethierryi Reiche, 1868
Brachinus lewecki Liebke, 1928
Brachinus lewisii Bates, 1873
Brachinus limbellus Chaudoir, 1876
Brachinus limbicollis Chaudoir, 1876
Brachinus limbiger Chaudoir, 1876
Brachinus lombokensis Kirschenhofer, 2010 ID
Brachinus longipalpis Wiedemann, 1821
Brachinus longulus Chaudoir, 1876
Brachinus luzonicus Chaudoir, 1876
Brachinus macrocerus Chaudoir, 1876
Brachinus mactus Peringuey, 1904
Brachinus madecassus (Jeannel, 1949)
Brachinus magyari Jedlicka, 1960
Brachinus marginellus Dejean, 1826
Brachinus marginipennis Motschulsky, 1864
Brachinus marginiventris Brulle, 1838
Brachinus marinus Motschulsky, 1845
Brachinus marleyi Barker, 1919
Brachinus maublanci (Colas, 1942) F
Brachinus mauretanicus Bedel, 1914
Brachinus medius Harris, 1828
Brachinus melanarthrus Chaudoir, 1876
Brachinus melancholicus Schmidt-Goebel, 1846
Brachinus merkli Kirschenhofer, 2003
Brachinus mesopotamicus Ali, 1967
Brachinus methneri Liebke, 1934
Brachinus mexicanus Dejean, 1831
Brachinus micheli (Jeannel, 1949)
Brachinus microamericanus Erwin, 1969
Brachinus microrrhabdus (Alluaud, 1899)
Brachinus midoli Alluaud, 1917
Brachinus minor Jedlicka, 1958
Brachinus mobilis Erwin, 1970
Brachinus modestus Schmidt-Goebel, 1846
Brachinus motschulskyi Jedlick
Brachinus muchei Jedlicka, 1967
Brachinus natalicus Peringuey, 1896
Brachinus neglectus Leconte, 1844
Brachinus niger Chaudoir, 1876
Brachinus nigricans Chaudoir, 1850
Brachinus nigricornis Gebler, 1829
Brachinus nigridorsis Nakane, 1962
Brachinus nigripes G.R.Waterhouse, 1841
Brachinus nigrovirescens Basilewsky, 1951
Brachinus nobilis Dejean, 1831
Brachinus nuristanus Jedlicka, 1967
Brachinus oaxacensis Erwin, 1970
Brachinus obliquetruncatus Peris, 1875
Brachinus obliterus Peringuey, 1896
Brachinus oblongus Dejean, 1825
Brachinus obscuripennis Motschulsky, 1864
Brachinus obscuritarsis Motschulsky, 1864
Brachinus obtusus (Thunberg, 1784)
Brachinus olgae Arribas, 1993
Brachinus olidus Reiche, 1842
Brachinus oneili Peringuey, 1899
Brachinus opacipennis Motschulsky, 1864
Brachinus orestes Kirschenhofer, 2003
Brachinus orestes Kirschenhofer, 2003
Brachinus orientalis Chaudoir, 1876
Brachinus ornatus Fairmaire, 1901
Brachinus otini Antoine, 1963
Brachinus ovipennis LeConte, 1862
Brachinus oxygonus Chaudoir, 1843
Brachinus pachygaster Perty, 1830
Brachinus pallidipes Reitter, 1919
Brachinus pallidus Erwin, 1965
Brachinus pallipes Dejean, 1826
Brachinus papua Darlington, 1968
Brachinus pateri Puel, 1938
Brachinus patruelis Leconte, 1844
Brachinus paviei Lesne, 1896
Brachinus pecoudi Puel, 1925
Brachinus pectoralis Dejean, 1825
Brachinus peltastes Andrewes, 1931
Brachinus perplexus Dejean, 1831
Brachinus perrieri (Jeannel, 1949)
Brachinus phaeocerus Chaudoir, 1868
Brachinus philippinensis Tian & Deuve, 2007
Brachinus piceus Chaudoir, 1876
Brachinus pictus Hope, 1833
Brachinus plagiatus Reiche, 1868
Brachinus planiusculus Motschulsky, 1864
Brachinus posticus Dejean, 1831
Brachinus praestans Andrewes, 1931
Brachinus promontorii Peringuey, 1888
Brachinus proximus Fairmaire, 1887
Brachinus pseudocruciatus Reitter, 1909
Brachinus psophia Audinet-Serville, 1821
Brachinus puberulus Chaudoir, 1868
Brachinus punctaticollis Heller, 1923
Brachinus puncticollis Schmidt-Goebel, 1846
Brachinus pygmaeus Dejean, 1826
Brachinus quadriguttatus Gebler, 1829
Brachinus quadripennis Dejean, 1825
Brachinus reyi Andrewes, 1924
Brachinus rikatlae Peringuey, 1896
Brachinus rugipennis Chaudoir, 1868
Brachinus sallei Chaudoir, 1876
Brachinus sanctus Motschulsky, 1864
Brachinus schmidti Andrewes, 1927
Brachinus scitulus Schmidt-Goebel, 1846
Brachinus sclopeta Fabricius, 1792
Brachinus scotomedes L. Redtenbacher, 1867
Brachinus scotti Liebke, 1934
Brachinus scriptus Chaudoir, 1878
Brachinus scutellatus Chaudoir, 1876
Brachinus sericeus Dejean, 1831
Brachinus servillei Marc, 1839
Brachinus sexmaculatus Dejean, 1825
Brachinus sexnotatus Liebke, 1934
Brachinus sexpustulatus (Fabricius, 1775)
Brachinus seyrigi (Alluaud, 1935)
Brachinus sibiricus Motschulsky, 1864
Brachinus sicardi Jeannel, 1949
Brachinus sichemita Reiche & Saulcy, 1855
Brachinus simulans Peringuey, 1896
Brachinus solidipalpis Tian & Deuve, 2007
Brachinus solidus Peringuey, 1899
Brachinus somereni Burgeon, 1947
Brachinus sonorous Erwin, 1970
Brachinus sordidus Andrewes, 1933
Brachinus stappersi Liebke, 1934
Brachinus stenoderus Bates, 1873
Brachinus stevensi Andrewes, 1924
Brachinus stevensianus Jedlicka, 1956
Brachinus strictus Morvan, 1971
Brachinus stygius Andrewes, 1933
Brachinus subcostatus Dejean, 1825
Brachinus suberbianus Alluaud, 1935
Brachinus sublaevis Chaudoir, 1868
Brachinus subnotatus Chaudoir, 1844
Brachinus suensoni Kirschenhofer, 1986
Brachinus sulcipennis (Jeannel, 1949)
Brachinus suturalis (Jeannel, 1949)
Brachinus suturatus Chaudoir, 1876
Brachinus suturellus Chaudoir, 1876
Brachinus szetschuanensis Jedlicka, 1963
Brachinus talyschensis Motschulsky, 1850
Brachinus tenuicollis LeConte, 1844
Brachinus tenuis Lorenz, 1998
Brachinus testaceus Rambur, 1837
Brachinus tetracolon Chaudoir, 1876
Brachinus tetragrammus Chaudoir, 1876
Brachinus tetraspilotus Chaudoir, 1876
Brachinus tetrastigma Fairmaire, 1897
Brachinus texanus Chaudoir, 1868
Brachinus thieleni Kirschenhofer, 1987
Brachinus tianshanicus Mikhailov, 1976
Brachinus tigridis Ali, 1967
Brachinus truncatulus Fairmaire, 1901
Brachinus tsara Alluaud, 1918
Brachinus turkestanicus Liebke, 1928
Brachinus vadoni Jeannel, 1949
Brachinus vagus Peringuey, 1899
Brachinus variegatus (Roth, 1851)
Brachinus variventris Schaufuss, 1862
Brachinus velutinus Erwin, 1965
Brachinus vicinus Dejean, 1826
Brachinus vigilans Chaudoir, 1876
Brachinus viridipennis Dejean, 1831
Brachinus vitticollis Chaudoir, 1876
Brachinus vulcanoides Erwin, 1969
Brachinus xanthophryus Chaudoir, 1876
Brachinus xanthopleurus Chaudoir, 1876
Brachinus yunnanus Jedlicka, 1963

Litteratur

Bertram G. Rye. 1908. Biller I. Løpebiller. Danmarks Fauna Bind 3. G.E.C. GAD, København.

Eksterne lenker

(en) Brachinus i Encyclopedia of Life
(en) Brachinus i Global Biodiversity Information Facility
(no) Brachinus hos Artsdatabanken
(sv) Brachinus hos Dyntaxa
(en) Brachinus hos Fauna Europaea
(en) Brachinus hos Fossilworks
(en) Brachinus hos ITIS
(en) Brachinus hos NCBI
(en) Kategori:Brachinus – bilder, video eller lyd på Wikimedia Commons
(en) Brachinus – galleri av bilder, video eller lyd på Wikimedia Commons
Brachinus – detaljert informasjon på Wikispecies
Tree of Life – Bombardérbiller (Brachininae, Carabidae) – engelskspråklig
Brachininae - fotogalleri - engelskspråklig
Norsk Entomologisk forening

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