Tachys

Tachys
	Tachys scutellaris
	Nomenklatur
	Tachys; Dejean, 1821
	Populærnavn
	skyggeløpere
	Klassifikasjon
Rike	Dyreriket
Rekke	Leddyr
Klasse	Insekter
Orden	Biller
Familie	Løpebiller
Underfamilie	Skyggeløpere
	Økologi
Antall arter:	ca. 300
Habitat:	terrestrisk, i stor grad underjordisk
	Inndelt i

Tachys er slekt av små løpebiller, gjerne rødbrune på farge.

Utseende[rediger | rediger kilde]

Små (gjerne 2-3 millimeter) løpebiller, kroppsformen forholdsvis slank og oftest litt flat, overflaten oftest blank. Mange er mer eller mindre rødbrune, men det finnes også mange blanksvarte eller metalliske arter. Hodet er ofte forholdsvis lite og smalt med store, fremstrakte overkjever (mandibler). Fasettøynene er middels store, litt utstående. Antennene er vanligvis lange og tynne. Det ytterste leddet på maksillarpalpene (munnfølere) er redusert til en nålefin tapp. Brystskjoldet (pronotum) er rundet, vanligvis med markerte bakhjørner. Dekkvingene er jevnbrede uten framtredende "skuldre", gjerne uten tydelige punktrader med med noen oppstående børster. Beina er nokså lange og slanke.

Levevis[rediger | rediger kilde]

Skyggeløperne kan finnes i mange ulike miljøer på bakken, og er ofte svært tallrike. De er rovdyr og angriper ulike små byttedyr. De fleste er raske løpere, men en god del arter har mistet flygeevnen.

Systematisk inndeling[rediger | rediger kilde]

Treliste

Ordenen biller, Coleoptera Linnaeus, 1758
- Underordenen Adephaga
  - Overfamilien løpebiller og vannkalver, Caraboidea
    - Familien løpebiller, Carabidae Latreille, 1802
      - Underfamilien skyggeløpere, Trechinae Bonelli, 1810
        Stammen Bembidiini Stephens, 1827
        Gruppen Tachyina Motschulsky, 1862
        Slekten Tachys Dejean, 1821

Liste over artene[rediger | rediger kilde]

Tachys abruptus Darlington, 1934^{c g}
Tachys aeneipennis Motschulsky, 1862^{i c g}
Tachys aequinoctialis (Motschulsky, 1855)^{c g}
Tachys albipes Leconte, 1863
Tachys algiricus (Lucas, 1846)^{c g}
Tachys ambulatus Darlington, 1962^{c g}
Tachys androyanus (Jeannel, 1946)^{c g}
Tachys angustulus Reitter, 1899^{c g}
Tachys apex Darlington, 1962^{c g}
Tachys arcanicola Blackburn, 1878^{i c g}
Tachys arculus Fauvel, 1882^{c g}
Tachys argentinicus Csiki, 1928^{c g}
Tachys artensis Montrouzier, 1860
Tachys asemus (Basilewsky, 1968)^{c g}
Tachys assamicus (Jedlicka, 1964)^{c g}
Tachys atamuradovi Kryzhanovskij & Mikhailov, 1987^{c g}
Tachys atomarius Wollaston, 1867^{c g}
Tachys atomus Blackburn, 1878^{i c g}
Tachys atratus A.Costa, 1883^{c g}
Tachys atridermis Sloane, 1921^{c g}
Tachys austinicus Casey, 1918
Tachys australicus Sloane, 1896^{c g}
Tachys avius Darlington, 1962^{c g}
Tachys baldiensis Blackburn, 1891^{c g}
Tachys barbieri Straneo, 1953^{c g}
Tachys bathyglyptus Andrewes, 1925^{c g}
Tachys beatus Darlington, 1962^{c g}
Tachys beaumonti Casey, 1918^{c g}
Tachys bicoloratus Burgeon, 1935^{c g}
Tachys bistriatus (Duftschmid, 1812)^{c g}
Tachys blackburni Sloane, 1921^{c g}
Tachys blemoides (Jeannel, 1946)^{c g}
Tachys bloetei Andrewes, 1930^{c g}
Tachys bolellus Darlington, 1962^{c g}
Tachys bolus Darlington, 1962^{c g}
Tachys bonariensis Steinheil, 1869^{c g}
Tachys borkuensis (Bruneau de Miré, 1952)^{c g}
Tachys bouchardi Andrewes, 1931^{c g}
Tachys brachys Andrewes, 1925^{c g}
Tachys bradycellinus Hayward, 1900^{i c g}
Tachys bredoi (Basilewsky, 1953)^{c g}
Tachys brevicornis Chaudoir, 1846
Tachys brunneus Andrewes, 1925^{c g}
Tachys brunniceps Andrewes, 1930^{c g}
Tachys bryanti Lindroth, 1966^{i c g}
Tachys caffer Péringuey, 1896^{c g}
Tachys caheni (Basilewsky, 1972)^{c g}
Tachys caledonicus Csiki, 1928^{c g}
Tachys callipygus (Boheman, 1858)^{c g}
Tachys cameroni Andrewes, 1925^{c g}
Tachys captus Blackburn, 1888^{c g}
Tachys cardioderus Chaudoir, 1850^{c g}
Tachys cardoni Andrewes, 1925^{c g}
Tachys carib Darlington, 1936^{c g}
Tachys carinulatus Sloane, 1921^{c g}
Tachys caspius Kolenati, 1845^{c g}
Tachys castaneicolor Bates, 1882^{c g}
Tachys centralis J.sahlberg, 1900
Tachys centriustatus Reitter, 1894
Tachys centromaculatus Wollaston, 1864^{c g}
Tachys chiriquinus Bates, 1882^{c g}
Tachys cinctus Putzeys, 1875^{c g}
Tachys collardi (Basilewsky, 1955)^{c g}
Tachys colonicus Casey, 1918^{c g}
Tachys columbiensis Hayward, 1900^{i c g}
Tachys corax Leconte, 1852
Tachys coriaceus Broun, 1908^{c g}
Tachys crypticola (Britton, 1960)^{c g}
Tachys cubax Darlington, 1934^{c g}
Tachys curtulus (Basilewsky, 1953)^{c g}
Tachys curvans Bates, 1882
Tachys cycloderus Bates, 1871^{c g}
Tachys cyprius Coulon, 2004^{c g}
Tachys delamarei (Jeannel, 1962)^{c g}
Tachys delicatus Andrewes, 1925^{c g}
Tachys dentatus Andrewes, 1925^{c g}
Tachys descarpentriesi (Bruneau de Miré, 1965)^{c g}
Tachys devroeyi Burgeon
Tachys diaphanus Casey, 1918
Tachys dimediatus Motschulsky, 1849^{c g}
Tachys dimidiatus Motschulsky, 1849^g
Tachys diminutus Bates, 1871^{c g}
Tachys diploharpinus Bates, 1878^{c g}
Tachys discipennis Fauvel, 1903
Tachys dives Péringuey, 1926^c
Tachys dominicanus Darlington, 1934^{c g}
Tachys dorsalis Motschulsky, 1851^{c g}
Tachys dostali Baehr, 2014^{c g}
Tachys dromioides Bates, 1871^{c g}
Tachys dzosonicus Pawlowski, 1974^{c g}
Tachys ectromioides Sloane, 1896^{c g}
Tachys edax Leconte, 1852
Tachys ellenbergeri (Bruneau de Miré, 1964)^{c g}
Tachys elongatulus (Dejean, 1831)^{c g}
Tachys enormis (Dostal, 2015)^{c g}
Tachys erwini Reichardt, 1976^{c g}
Tachys euryodes Bates, 1892^{c g}
Tachys excisicollis Baehr, 2012^{c g}
Tachys exochrias Darlington, 1962^{c g}
Tachys fasciatus (Motschulsky, 1851)^{c g}
Tachys federicae Baehr, 2016^{c g}
Tachys filax Darlington, 1934^{c g}
Tachys flavax Darlington, 1962^{c g}
Tachys flavicollis Motschulsky, 1862^{c g}
Tachys fortestriatus Baehr, 2003^{c g}
Tachys fortunatus Machado, 1989^{c g}
Tachys frischi Coulon & Felix, 2009
Tachys fulvicollis (Dejean, 1831)^{c g}
Tachys fumax Darlington, 1962^{c g}
Tachys gilvus Schaum, 1863^{c g}
Tachys gracchus Andrewes, 1935^{c g}
Tachys gratus Péringuey, 1896^c
Tachys guangxicus (Deuve & Tian, 2010)^{c g}
Tachys gyotokuensis Tanaka, 1956^{c g}
Tachys halophilus Lindroth, 1966^{i c g}
Tachys haywardi Wickham, 1913^{c g}
Tachys holmi (Basilewsky, 1963)^{c g}
Tachys hyalinus Casey, 1918^{i c g}
Tachys hydrophilus (Germain, 1906)^{c g}
Tachys impictus Andrewes, 1925^{c g}
Tachys impressipennis Motschulsky, 1859^{c g}
Tachys impressus Motschulsky, 1851^{c g}
Tachys incertus Andrewes, 1925^{c g}
Tachys infuscatus Blackburn, 1888^{c g}
Tachys intermedius Baehr, 2012^{c g}
Tachys iridipennis Chaudoir, 1876^{c g}
Tachys jeannei Coulon, 2004^{c g}
Tachys joannae (Basilewsky, 1962)^{c g}
Tachys jucundulus Péringuey, 1908^{c g}
Tachys kabylianus Puel, 1935^{c g}
Tachys kahuzianus (Basilewsky, 1953)^{c g}
Tachys kaorutanakai Habu, 1977^{c g}
Tachys koizumii Habu, 1961^{c g}
Tachys koreanorum Pawlowski, 1974^{c g}
Tachys laevigatus (Boheman, 1858)^{c g}
Tachys laevus (Say, 1823)^{c g}
Tachys lamottei (Bruneau de Miré, 1964)^{c g}
Tachys languidus Andrewes, 1925^{c g}
Tachys latalatus Csiki, 1928^{c g}
Tachys leleupi (Basilewsky, 1953)^{c g}
Tachys lenkoranus Csiki, 1928^{c g}
Tachys leptocerus Chaudoir, 1876^{c g}
Tachys leytensis (Baehr, 2016)^{c g}
Tachys limbatellus Bates, 1884^{c g}
Tachys lindi Blackburn, 1888^{c g}
Tachys lissonotus Andrewes, 1925^{c g}
Tachys litoralis Casey, 1884^{i c g}
Tachys lividus (Bates, 1871)^{c g}
Tachys longulus Andrewes, 1925^{c g}
Tachys lugubris Motschulsky, 1862^{c g}
Tachys lusciosus Antoine, 1943
Tachys luscus Darlington, 1962^{c g}
Tachys macleayi Sloane, 1896^{c g}
Tachys macrops Baehr, 2012^{c g}
Tachys mameti Alluaud, 1933^{c g}
Tachys marggii Kirschenhofer, 1986^{c g}
Tachys marri Baehr, 1987^{c g}
Tachys masculus Darlington, 1962^{c g}
Tachys massauxi (Basilewsky, 1955)^{c g}
Tachys mastersi Sloane, 1921^{c g}
Tachys micros (Fischer von Waldheim, 1828)^{c g}
Tachys microscopicus (Bates, 1873)^{c g}
Tachys minusculus (Motschulsky, 1862)
Tachys minutissimus (R.F.Sahlberg, 1847)^c
Tachys mirandus Coulon & Wrase, 2009
Tachys mirei (Basilewsky, 1968)^{c g}
Tachys misellus Laferte-Senectere, 1841
Tachys mitchelli Sloane, 1895^{c g}
Tachys moira (Erwin, 1984)
Tachys monostictus Bates, 1871^{c g}
Tachys mordax Leconte, 1852
Tachys mulwalensis Sloane, 1900^{c g}
Tachys mus Andrewes, 1925^{c g}
Tachys myrmecophilus (Jeannel, 1946)^{c g}
Tachys nanniscus Péringuey, 1896^{c g}
Tachys naraensis Ueno, 1953^{c g}
Tachys nephelodes Andrewes, 1930^{c g}
Tachys niger (Jeannel, 1953)^{c g}
Tachys nitmiluki (Giachino, 2003)^{c g}
Tachys nympha (Erwin, 1984)
Tachys oahuensis Blackburn, 1878^{i c g}
Tachys oblitus Casey, 1918^{i c g b}
Tachys obtusiusculus (Jeannel, 1941)^{c g}
Tachys ochrias Andrewes, 1925^{c g}
Tachys ochrioides Darlington, 1962^{c g}
Tachys olemartini Kirschenhofer, 1986^{c g}
Tachys opalescens Andrewes, 1925^{c g}
Tachys orphninus Andrewes, 1925^{c g}
Tachys otini (Antoine, 1944)
Tachys pallidus Chaudoir, 1868^{i c g b}
Tachys pallorus Kopecky in Löbl & Smetana, 2003^{c g}
Tachys palustris Reitter, 1894
Tachys panamensis Casey, 1918^{c g}
Tachys particula Andrewes, 1936^{c g}
Tachys paulax Darlington, 1934^{c g}
Tachys pharao Schatzmayr & Koch, 1934^{c g}
Tachys piceolus Laferte-Senectere, 1841
Tachys plagiatus Putzeys, 1875^{c g}
Tachys pliginskii Solodovnikov, 2001^{c g}
Tachys politissimus Baehr, 2014^{c g}
Tachys potomaca (Erwin, 1981)^{i b}
Tachys potomacus (Erwin, 1981)^{c g}
Tachys prionotus Andrewes, 1936^{c g}
Tachys privus Darlington, 1962^{c g}
Tachys prolixus Bates, 1892^{c g}
Tachys proximus Say, 1823
Tachys pseudolusciosus Coulon, 2004^{c g}
Tachys pseudosericeus Kirschenhofer, 1986^{c g}
Tachys pujoli (Bruneau de Miré, 1964)^{c g}
Tachys pulchellus Laferte-Senectere, 1841
Tachys pumilus (Dejean, 1831)^{i c g b}
Tachys pusillimus Péringuey, 1896^{c g}
Tachys quadrillum Schaum, 1860
Tachys queenslandicus Sloane, 1903^{c g}
Tachys rambai Jedlicka, 1966^{c g}
Tachys rapoporti (Jeannel, 1962)
Tachys rectangulus Notman, 1919^{i c g}
Tachys rhodeanus Casey, 1918^{i c g b}
Tachys ridiculus (L.Schaufuss, 1879)
Tachys riedeli Baehr, 2014^{c g}
Tachys rufotestaceus Hayward, 1900
Tachys sagax Casey, 1918^{i c g}
Tachys schuberti Jedlicka, 1968^{c g}
Tachys scitulus Leconte, 1848^{i c g b}
Tachys scutellaris Stephens, 1828^{c g}
Tachys sellatus Fairmaire, 1892^{c g}
Tachys sequax Leconte, 1848
Tachys sericans Bates, 1873^{c g}
Tachys sericeus Motschulsky, 1851^{c g}
Tachys serra Darlington, 1962^{c g}
Tachys serratus Andrewes, 1925^{c g}
Tachys serrula Darlington, 1962^{c g}
Tachys sexguttatus Fairmaire, 1849
Tachys shahinei Schatzmayr & Koch, 1934^{c g}
Tachys shirazi Jedlicka, 1968^{c g}
Tachys sibling Darlington, 1962^{c g}
Tachys similis Blackburn, 1888^{c g}
Tachys simulator Coulon, 2004^{c g}
Tachys spadix Casey, 1918^{i c g}
Tachys striax Darlington, 1934^{c g}
Tachys subangulatus Bates, 1871^{c g}
Tachys subbrunneus Darlington, 1962^{c g}
Tachys sublobatus Darlington, 1962^{c g}
Tachys suboculatus (Basilewsky, 1958)^{c g}
Tachys sudanicus (Basilewsky, 1948)^{c g}
Tachys suensoni Kirschenhofer, 1986^{c g}
Tachys sundaicus Andrewes, 1925^{c g}
Tachys suturifer Reitter, 1884^{c g}
Tachys sydneyensis Sloane, 1923^{c g}
Tachys szekessyi Jedlicka, 1968^{c g}
Tachys tantillus Motschulsky, 1862
Tachys tenuiserra Darlington, 1962^{c g}
Tachys terra (Erwin, 1984)
Tachys testaceus (Basilewsky, 1953)^c
Tachys tetraphacus Bedel, 1896^{c g}
Tachys tienmushaniensis Kirschenhofer, 1986^{c g}
Tachys torretassoi Schatzmayr & Koch, 1934^{c g}
Tachys transcaucasicus Coulon, 2004^{c g}
Tachys translucens Darlington, 1937^{i c g}
Tachys transumbratus Bates, 1892^{c g}
Tachys transversicollis (W.J.Macleay, 1871)
Tachys trechoderus Coulon, 2004^{c g}
Tachys trechulus Darlington, 1936^{c g}
Tachys troglocola (Deuve & Tian, 2010)^{c g}
Tachys troglophilus Ueno, 1953^{c g}
Tachys tropicalis Baehr, 2012^{c g}
Tachys tropicus (Nietner, 1858)^{c g}
Tachys truncatus (Nietner, 1858)^{c g}
Tachys turkestanicus Csiki, 1928
Tachys ucayali (Erwin, 1984)
Tachys uelensis Burgeon, 1935^{c g}
Tachys uenoianus Habu, 1974^{c g}
Tachys umbripennis Chaudoir, 1868^{i c g}
Tachys uniformis Blackburn, 1888^{c g}
Tachys vandeli (Mateu & Colas, 1954)^{c g}
Tachys vandepolli (Bruneau de Miré, 1964)^{c g}
Tachys varsavianorum Pawlowski, 1974^{c g}
Tachys ventricosus Leconte, 1863
Tachys venustus Andrewes, 1936^{c g}
Tachys vernilis Casey, 1918^{i c g}
Tachys vibex Kopecky in Löbl & Smetana, 2003^{c g}
Tachys vietnami Jedlicka, 1968^{c g}
Tachys vilis Andrewes, 1925^{c g}
Tachys virgo Leconte, 1852
Tachys vittiger Leconte, 1852
Tachys vorax Leconte, 1852
Tachys windsorensis Baehr, 1991^{c g}
Tachys xanthochrous Chaudoir, 1876^{c g}
Tachys yeboensis Burgeon, 1935^{c g}
Tachys yodai Jedlicka, 1964^{c g}
Tachys yorkensis Baehr, 2012^{c g}
Tachys yunchengensis Kirschenhofer, 1986^{c g}
Tachys zonatus Andrewes, 1925^{c g}
Tachys zulficari Schatzmayr & Koch, 1934^{c g}

Litteratur[rediger | rediger kilde]

Eksterne lenker[rediger | rediger kilde]

(en) Tachys – oversikt og omtale av artene i WORMS-databasen
(en) Tachys i Encyclopedia of Life
(en) Tachys i Global Biodiversity Information Facility
(no) Tachys hos Artsdatabanken
(sv) Tachys hos Dyntaxa
(en) Tachys hos Fauna Europaea
(en) Tachys hos Fossilworks
(en) Tachys hos ITIS
(en) Tachys hos NCBI
(en) Kategori:Tachys – bilder, video eller lyd på Wikimedia Commons
Tachys – detaljert informasjon på Wikispecies
Bilder: